Doug came to Janelia Farm from Johns Hopkins Medical School in Baltimore where he was a professor of cell biology for 30 years and organized a core facility in microscopy in the medical school. Doug's research lab investigated the role of microtubules in organelle transport and cell motility; microtubule polymer dynamics; and the functions of microtubule-associated proteins such as MAP-2 in neurons and other cell types. Doug received his Ph.D. from the University of Pennsylvania, and also studied at the University of Rochester (B.A.), Syracuse (M.S.), and the University of Wisconsin (postdoc). While at Hopkins, Doug received a Senior Fulbright Lectureship to teach in cell biology at Moscow University and participated in National Academy of Sciences visitor exchanges to do research in the USSR. In 2001, Doug wrote a textbook entitled Fundamentals of Light Microscopy and Electronic Imaging and more recently has contributed review articles and Java tutorials to the Olympus optical microscopy educational websites.